中国及泛喜马拉雅地区草糙苏属（唇形科）的分类学研究

The genus Phlomoides Moench was resurrected and redefined to include five genera: Eremostachys Bunge, Lamiophlomis Kudô, Notochaete Benth., Phlomis s. str., and Pseuderemostachys Popov. As currently defined, Phlomoides consists of ca. 150 species and is universally recognized as a taxonomically difficult genus of Lamiaceae. The Pan-Himalaya regions (including SW China) is the center of distribution and diversity of the genus, of which most are endemic species, and some of them are economically and medicinally important. Although there are a lot of taxonomic problems need to be resolved and lack of firsthand information, the genus appeals low attention from botanist and the main reason is that the genus was recognized as a distinct genus until recently. .Based on extensive field work and herbarium specimens’ investigation, as well as cytological and morphological studies, this study will resolve these taxonomic problems for the genus: 1, reveal species boundaries and clarify the existed taxonomic confusion; 2, clarify the species diversity of the genus in China and the Pan-Himalayas, and outline distribution area for each species; 3, propose a reasonable and reliable infrageneric classification of the genus. .This study could be the final piece of the jigsaw to accomplish the volume Lamiaceae of “Flora of Pan-Himalayas”, and will lay a foundation for worldwide taxonomic review of the genus in future studies. Meanwhile, this study will promote the taxonomic and systematic study of the subfamily Lamioideae, and commit to personnel training for classic taxonomy for such specific large group.

草糙苏属为新近恢复的大属，且属的范围发生了重要变化，包括了其他5个属的物种。该属是唇形科植物分类研究中尤其困难的类群之一，共约150种。泛喜马拉雅地区（含我国西南大部分地区）为其分布中心和多样化中心，特有种多，很多物种具有重要的药用和观赏价值。因草糙苏属的属级分类地位最近才被认可，虽然存在的分类学问题较多，但关注度却很低，研究资料也极度匮乏。. 本项目拟通过野外考察和标本查阅，结合细胞学、形态解剖学等研究资料，解决该属存在的以下分类学问题：1，明确物种界限，澄清分类疑难；2，确定中国及泛喜马拉雅地区的物种多样性及各物种的地理分布情况；3，提出一个合理的属下分类系统。. 本项目不仅是《泛喜马拉雅植物志》唇形科编研的最后一块拼图，也将为后期完成该属世界范围的分类修订奠定基础，在促进野芝麻亚科的分类学和系统学研究的同时，还为培养专科专属的植物经典分类人才做出贡献。

草糙苏属为新近恢复的大属，且属的范围发生了重要变化，包括了其他5个属的物种。该属是唇形科植物分类研究中尤其困难的类群之一，共约150种。泛喜马拉雅地区（含我国西南大部分地区）为其分布中心和多样化中心，特有种多，很多物种具有重要的药用和观赏价值。草糙苏属的属级分类地位最近才被认可，虽然存在的分类学问题较多，但关注度却很低，研究资料也极度匮乏。.本项目针对中国及泛喜马拉雅地区该属物种存在的分类学问题，开展了全面的分类学和分子系统学研究工作，查阅国内外22个标本馆的馆藏标本6000余份（模式标本350份），收集分子材料227号137种，采集植物标本308号。基于119个样品的分子系统学研究结果，进一步明确了草糙苏属的界限，将箭叶水苏属、短唇沙穗属并入草糙苏属。完成了47种6变种共80份的苞片电镜扫描以及48种6变种共88份叶片的显微形观察，以及该属29种3变种共39份的小坚果微形态观察和电镜扫描，明确了基生叶、苞叶、苞片、花萼、小坚果、雄蕊后对花丝基部附属器的形态多样性与变异规律，揭示了这些重要性状的分类学价值。最终，结合多学科研究证据，明确了该地区共有草糙苏属植物57种，发现了2个新种和1个中国新记录种，提出了16个新异名和5个新组合，并编制了物种检索表。.依托于该项目的实施，共发表科研论文8篇，培养硕士研究生1人，团队成员中有两人分别获得了博士和硕士学位，为培养专科专属的植物经典分类人才做出贡献。