华南二叠纪至三叠纪海洋无脊椎动物之间捕食作用与共生关系的演化过程与模式

This new project aims to systematically study the Late Permian to Middle Triassic paleontology and paleoecology of 20 sections, which represent various depsotional environments ranging from intraplatform basin, ramp, shallow platform, siliciclastic shallow sea, to nearshore settings in South China. Emphasized are these fossil records showing predation and symbiotic interactions among marine invertebrates. Detailed morphologic classification, measurement and quantitative analysis will be conducted on the predatory drillholes, bite marks, and shell-crushing marks as well as episkeletobionts and endoskeletobionts. This study also attempts to reveal the predator-prey pressure and skeletobiont-host competition in ultilizing ecologic resources, forawrd the replacement models of predation and symbiotic interactions in various environmental settings during biotic extinction, survival and recovery stages, with emphasis on evolutionary traits of biotic predation pressure variations in marine ecosystems before and after the mass extinction as well as during the biotic recovery. This new project also hopes evoluate potential warning signals of biotic predation and symbiosis response to biotic mass extinction and possible driving dynamic faciliating ecosystem's restoration. Eventually, the global models of predation and symbiosis evolutions of invertebrates during this critical period of macroevolution of Earth life will be established, providing some examples for unravelling the organism-organism and organism-environment co-evolutions during major geological turning periods.

新项目拟对华南地区20条代表不同水深环境剖面的上二叠统至中三叠统地层进行深入的古生物学和古生态学研究，强调地层中保存的古无脊椎动物之间的捕食作用和共生关系化石记录，特别是对化石中记录的各类钻孔、咬痕、破碎壳等捕食作用证据加以生态特征分析、进行度量、统计和定量分析，评价捕食者-猎物之间生存压力。同时，对各类以介壳为基底的表生和内生共生生物进行分类和定量生态学分析，揭示共生生物与寄宿主之间竞争资源的关系以及外部环境对它们的影响机制，建立不同环境中生物之间在大灭绝前、后以及大灾难之后生态系统复苏重建过程中捕食作用和共生关系的更替模式，评价生物捕食作用和共生关系对生态系统坍塌潜在的预警能力及灾后生态系统重建可能的驱动机制。最终，建立该时期全球生物捕食与共生关系演化模式，并为揭示重大地质突变期生物与生物之间以及生物与环境的协同演化提供实证。

为揭示重大地质突变期生物与生物之间以及生物与环境的协同演化提供实证，本项目对华南地区不同水深环境剖面的上二叠统至中三叠统地层进行深入的古生物学和古生态学研究，对化石中记录的各类钻孔、咬痕、破碎壳等捕食作用证据加以生态特征分析、进行度量、统计和定量分析，评价捕食者-猎物之间生存压力。对各类以介壳为基底的表生和内生共生生物进行分类和定量生态学分析，揭示共生生物与寄宿主之间竞争资源的关系以及外部环境对它们的影响机制，建立不同环境中生物之间在大灭绝前、后以及大灾难之后生态系统复苏重建过程中捕食作用和共生关系的更替模式，评价生物捕食作用和共生关系对生态系统坍塌潜在的预警能力及灾后生态系统重建可能的驱动机制。建立该时期全球生物捕食与共生关系演化模式。完成项目计划工作同时，以项目为依托完成包括Earth-Science Reviews，Geology，The Geological Society of America Bulletin等杂志SCI论文21篇。